Survival of the fitting

From Images of Organization:

Many biologists now believe that it is the whole ecosystem that evolves and the process of evolution can really be understood only at the level of the total ecology... it suggests that evolution is always evolution of a pattern of relations embracing organisms and their environments. It is the pattern, not just the separate units comprising this pattern, that evolves. Or as Kenneth Boulding has put it, evolution involves the "survival of the fitting" not just survival of the fittest.

Organizations and their environment are engaged in a pattern of cocreation, where each produces the other.